Evolutionary ecology: foraging, sex allocation, sex ratio, sexual selection & life-history. Metabolic ecology: body size, temperature & biological rates. Scaling & allometry in population biology.
- PhD, with Gordon Orians & Douglas Chapman(1973), Quantitative Ecology & Resource Management, University of Washington, Seattle
- Post Doctoral Fellow, with CS Holling, Institute of Animal Resource Ecology, University of British Columbia
Distinguished Professor of Biology, University of New Mexico