• Professor, Biology


  • Villa, S. M & J. A. H. Koop, C. Le Bohec and D. H. Clayton (2018). Beak of the pinch: Anti-parasite traits are similar among Darwin’s finch species. Evolutionary Ecology. Vol. 32, 443-452. Published, 08/2018.
  • Bush, S.E. & D. R. Gustafsson and D. H. Clayton (2018). New records of ectoparasites from passerine birds in the High Tatras of Slovakia. Oecologia Montana. Vol. 27, 43-45. Published, 07/2018.
  • Bush, S.E. (2018). Anti-parasite behavior of birds. Philosophical Transactions of the Royal Society of London B. Published, 06/2018.
  • DiBlasi, E.K. & K. P. Johnson, S.A. Stringham, A.N. Hansen, A.B. Beach, D.H. Clayton and S.E. Bush (2018). Phoretic dispersal influences parasite population genetic structure. Molecular Ecology. Published, 05/2018.
  • Villa, S. M & J. D. Evans, Y. K. Subhani, J. C. Altuna, S. E. Bush and D. H. Clayton (2018). Body size and fecundity are correlated in feather lice (Phthiraptera: Ischnocera): Implications for Harrison’s Rule. Ecol. Entomol. Published, 03/2018.
  • Gustafsson, D.R. & D. H. Clayton, and S. E. Bush (2018). Twelve new species of Priceiella Gustafsson & Bush, 2017 (Phthiraptera: Ischnocera: Philopteridae) from Old World babblers, with a key and comprehensive host-parasite checklist. Zootaxa. Vol. 4382, 401-449. Published, 02/2018.
  • McNew, S.M. & D. H. Clayton (2018). Alien invasion: Biology of Philornis flies highlighting P. downsi, an introduced parasite of Galapagos birds. Annual Review of Entomology. Vol. 63, 369-387. Published, 01/2018.
  • Boyd, B. M. (2017). Phylogenomics using target restricted assembly resolves intra-generic relationships of parasitic lice (Phthiraptera: Columbicola). Systematic Biology. Published, 11/17/2017.
  • McNew, S. M. (2017). Alien invasion: Biology of Philornis flies highlighting P. downsi, an introduced parasite of Galapagos birds. Annual Review of Entomology. Published, 10/20/2017.
  • Heimpel, G. E (2017). Invasive parasites and the fate of Darwin’s finches in the Galapagos Islands: the case of the Vegetarian Finch. Wilson Journal of Ornithology. Vol. 129, 345-349. Published, 09/17/2017.
  • McNew, S. M. (2017). Epigenetic variation between urban and rural populations of Darwin’s finches. BMC Evolutionary Biology. Published, 08/24/2017.
  • Knutie, S. A. (2017). Tri-trophic ecology of native parasitic nest flies of birds in Tobago. Ecosphere-Open Access Journal. Published, 01/27/2017.
  • Does allopreening control avian ectoparasites? Biology Letters 12: 20160362. http://dx.doi.org/10.1098/rsbl.2016.0362. Published, 07/26/2016.
  • Galápagos mockingbirds tolerant introduced parasites that threaten Darwin’s finches. Ecology 97:940-950. Published, 04/08/2016.
  • Does anti-parasite behavior improve with experience? Experimental test of the priming hypothesis. Behavioural Ecology doi: 10.1093/beheco/arw032. Published, 03/15/2016.
  • Clayton, D. H., S. E. Bush and K. P. Johnson. 2016. Coevolution of life on hosts: Integrating ecology and history. University of Chicago Press. 320 pages, 16 color plates, 110 halftones, 3 line drawings, 4 tables. Published, 12/2015.
  • Koop, J. A. H., P. S. Kim, S. A. Knutie, F. Adler and D. H. Clayton. 2015. Introduced parasitic fly may lead to local extinction of Darwin’s finch populations. Journal of Applied Ecology. doi: 10.1111/1365-2664.12575 Press coverage: Covered by BBC News, Smithsonian, Christian Science Monitor, Huffington Post, DailyMail, Fusion, MentalFloss and many others. Published, 06/2015.
  • Skinner, M. A., C. Gurerrero-Bosagna, Md M. Haque, E. E. Nilsson, J. A. H. Koop, Sarah A. Knutie and D. H. Clayton. 2014. Epigenetics and the evolution of Darwin's finches. Genome Biology and Evolution 6: 1972-1989. Published, 07/24/2014.
  • J.P. Owen, J. L. Waite, K. Z. Holden and D.H. Clayton. 2014. Does antibody binding to novel proteins predict future infection? (Invited article for special issue concerning immunology and ectoparasites). Parasite Immunology 36: 571-582. Published, 06/03/2014.
  • Ghosh, S., J.L. Waite, D. H. Clayton and F.R. Adler. 2014. Can antibodies against flies alter malaria transmission in birds by changing vector behavior? Journal of Theoretical Biology 358: 93-101. Published, 05/29/2014.
  • Knutie, S. A., S. M. McNew, A. W. Bartlow, D. Vargas, and D. H. Clayton. 2014. Darwin’s finches combat introduced nest parasites with fumigated cotton. Current Biology 24: R355–R356. Recommended by Faculty of 1000 Press coverage: Featured by Reuters, Science, Nature, National Public Radio, Scientific American, National Geographic, CBC Quirks & Quarks & many others. Published, 05/06/2014.
  • Waite, J.L., A.R. Henry, J.P. Owen and D.H. Clayton. 2014. An experimental test of the effects of behavioral and immunological defenses against vectors: do they interact to protect birds from blood parasites? Parasites & Vectors 7:104. Published, 03/04/2014.
  • Diversity of feather mites (Acari: Astigmata) on Darwin's Finches. J. Parasitology 99: 756-762. Published, 06/01/2013.


  • International Ornithological Congress, Vancouver, Canada. Invited Talk/Keynote, Presented, 08/2018.
  • 6th International Congress on Phthiraptera, Brno, Czech Republic. Contributed Talk, Presented, 06/2018.
  • Society for the Study of Evolution, Portland OR. Presentation, Presented, 10/20/2017.
  • American Museum of Natural History, New York. Invited Talk/Keynote, Presented, 08/24/2017.
  • Museum of Vertebrate Zoology, University of California, Berkeley. Invited Talk/Keynote, Presented, 06/20/2017.
  • Plenary lecture, Entomological Society of Canada Annual Meeting, Winnipeg. Invited Talk/Keynote, Presented, 05/10/2017.
  • Illinois Natural History Survey, University of Illinois, Urbana. Invited Talk/Keynote, Presented, 04/01/2017.
  • Douglas Schemske Retirement Symposium, University of Illinois, Chicago. Invited Talk/Keynote, Presented, 03/24/2017.
  • North American Ornithological Conference, Washington D. C. Contributed Talk, Presented, 08/15/2016.
  • Ecology and Evolutionary Biology class, University of Connecticut (Skype). Presentation, Presented, 08/01/2016.
  • Centre Méditerranéen Environnement et Biodiversité, Montpellier, France. Invited Talk/Keynote, Presented, 07/01/2016.
  • Society for the Study of Evolution, Austin TX . Contributed Talk, Presented, 05/01/2016.
  • Larada Sciences, 4rd Service Provider Summit, Park City, Utah . Invited Talk/Keynote, Presented, 04/01/2016.
  • Institute for High Mountain Biology, Žilina University, Slovak Republic. Invited Talk/Keynote, Presented, 06/2015.
  • Hungarian Natural History Museum, Budapest. Invited Talk/Keynote, Presented, 06/2015.
  • Dept. of Biology, University of Tulsa, Oklahoma. Invited Talk/Keynote, Presented, 05/2015.
  • Larada Sciences, 3rd Service Provider Summit, Park City, Utah. Invited Talk/Keynote, Presented, 04/2015.
  • Galápagos symposium, AAAS Pacific Division, San Francisco (Knutie). Invited Talk/Keynote, Presented, 04/2015.
  • Departmental Seminar, Dept. of Biology, University of Utah, Salt Lake City. Invited Talk/Keynote, Presented, 03/2015.
  • Microbial Pathogenesis Seminar Series, University of Utah, Salt Lake City. Invited Talk/Keynote, Presented, 03/2015.
  • Arthropods & Wildlife symposium, Entomol. Soc. America, Minneapolis (Knutie). Presentation, Presented, 01/2015.